Mutations

Mutations are the primary genetic layer. Each creature rolls exactly one. Hoard Gorgeous Fruit until you have an S-tier or SSS-tier monster worth re-rolling.

SStandard

Brave

Brave applies a +100% HP modifier. Dark Hiller with Brave eclipses 26,000 HP on a full optimized account.

RolePure Tank
TypeMutation
ObtainWild roll or Gorgeous Fruit
Best ForMain Tank - doubles HP
+100% HPPure Aggro SpongeTank Synergy
SStandard

Jolly

Jolly grants +80% HP plus a 15% HP regen on active skill use. The sustain slot for any long fight.

RoleSustain Tank
TypeMutation
ObtainWild roll or Gorgeous Fruit
Best ForSustain tanks / paladin archetype
+80% HP15% HP Regen on SkillLong-Fit Sustain
SSecret

Hasty

Hasty is the secret DPS mutation. +40% damage plus a 2.0x multiplier on undocumented advanced effects.

RolePrimary DPS
TypeMutation
ObtainWild roll or Gorgeous Fruit (secret drop)
Best ForHigh-velocity primary DPS with +40% damage and 2.0x advanced benefits
+40% Damage2.0x Advanced BenefitsDPS Sync
SSecret

Timid

Timid applies +40% damage per active party member AND reduces Hatred by 1. The ultimate backline DPS mutation.

RoleBackline DPS
TypeMutation
ObtainWild roll or Gorgeous Fruit (secret drop)
Best ForBackline artillery / glass cannon DPS
+40% Damage per Party Member-1 HatredBackline Anchor
SSecret

Naive

Naive applies a 3.0x multiplier to whatever Trait bonus is active. Pairs with Precision or Peerless for stacking value.

RoleSynergy Engine
TypeMutation
ObtainWild roll or Gorgeous Fruit (secret drop)
Best For3.0x multiplier on active Trait bonus (hybrid flex)
3.0x Trait MultiplierHybrid FlexCombo Engine
A+Standard

Naughty

Naughty is the consistent DPS mutation. Strong physical and magical damage scaling.

RoleStandard DPS
TypeMutation
ObtainWild roll or Gorgeous Fruit
Best ForReliable damage scaling on Milofoy / Dread Spike
Physical Damage BoostMagical Damage BoostConsistent DPS

Data Quality Notes

Keep entries useful by separating source-backed facts, player-tested strategy, and update timing.

Source fields

Keep rarity, obtain method, abilities, and summary tied to official posts or repeatable in-game testing.

Internal links

Link mutations entries to matching builds, boss routes, and guide sections.

Update rhythm

Keep tier values and obtain methods current after each game update.